Toadstool Fly
S1: anterior
Adelaide Hills
Red Eyed Lauxaniid Fly
Dorsal & Wing venation
 
    
Toadstool Fly (Tapeigaster paramonovi)Class: Animals (Animalia) - Jointed Legs (Arthropoda) - Insects (Insecta)
Order: Flies (Diptera)
Family: Toadstool Fly (Heteomyzidae)
Species: Toadstool Fly (Tapeigaster paramonovi)
This Photo:     S1: face

Thank you Tony Daley for identifying this species for us

General Species Information:
Found in the Adelaide Hills and possibly elsewhere
The mushroom S1 was sitting was measured later and estimate it's length to be ~6mm.
When we id'ed S1 as this species, we thought S2 was T. digitata but Tony kindly reviewed our findings and determined they were both the T. paramonovi. What is telling is that we thought S1 & S2 were the same fly, taken within 20 sec of each other. Now that is still possible and the differences showing are camera artifacts.
We mention these errors & our reasons below to highlight the difficulty in id'ing inverts to species and how expert opinion really is necessary with most inverts to get to species level.
The patch around the ocelli seems different. We noticed, however, there are many areas covered in very fine reflective felt/hair. As such, camera angle shows different colours. McAlpine & Kent's paper from 1982 states that T. digitata & T. paramonovi are "Closely related":
T. paramonovi has "whitish pruinescence along orbital margins"
T. digitata has "ocellar spot black"
There are also other differences related to leg colour, etc.
Tony said " In McAlpine's key (1982) T. digitata is split from the former species by the lack of mesonotum stripes and all tibiae with a black band above the middle and at apex. In the fly here, and as described for T. paramonovi, the fore tibiae has (only) the apical half or so darkened which appears to be unique among the genus, ie from middle to apex dark. My interpretation of ocelli spot colour for T. paramonovi from McAlpine's description is that it is also blackish by the lack of mention of colour when comparing to T. digitata, but instead differs from the latter by having it more or less dusted with whitish pruinescence."

Copyright © 2018- Brett & Marie Smith. All Rights Reserved. Photographed 02-Jun-2018
This species is an Australian Native Species, not listed in the SA Murray Mallee Survey of 2010.